Features
When it was known as the Harley Street of ceylon
The Homes in Ward Place in its early days,
(Continued from 16 May)
by Hugh Karunanayake, Dr Srilal Fernando, and Avinder Paul
The large four-acre property with the name Tyaganivasam (previously named Jaffna House) was the home of J Tyagarajah, member of the Monetary Board, and son of Namasivam Mudaliyar Tyagarajah. The grounds of Tyaganivasam included the property on which Cargills Pharmacy stood. Tyagarajah was also a Director of the Central Bank. He served in this capacity for more than two decades, never failing to attend meetings of the Monetary Board, and is reputed to have not claimed a cent for the expenditure incurred by him, a remarkable example of service to the nation. Part of the Tyagarajah property is now home to the University Grants commission.
With two major hospitals in close proximity, and despite the presence of Cargills Pharmacy at the opposite end of De Soysa Circus, the need for a pharmaceutical outlet in Ward Place was almost a sine qua non. The void was filled by the opening of the Lanka Pharmacy at 6, Ward Place by David Silva, who named it after his
son Lanka Silva, who stepped into the father’s shoes on leaving school. Lanka Silva was a champion athlete at Royal College of the early 1950s. “Manohari” The impressive home of Sir Arthur M De Silva ENT Surgeon was located nearby. His daughter married Justin Kotelawela, brother of former Prime Minister Sir John Kotelawela, in 1948.
Proceeding on the same side of Ward Place at No 16 stood Veerin the two storied home of Dr LAP Britto Babapulle a leading Veterinary surgeon of the time. Dr Babapulle was known as the owner of the largest number of tenement houses in Colombo, mostly located in the Grandpass area. His daughter, Andrea, lives in the house today.
A few properties away is Sukasthan Gardens, a cluster of homes built on the grounds of the former stately home of Sir Ponnambalam Ramanathan named “Sukasthan”. It was inherited by Ramanathan’s daughter, Sundari, who eventually sold it. Gynaecologist Dr PR Thiagarajah lived in one of the houses that were built there. Another well known resident of Sukasthan Gardens was LS Boys, a Director of Gordon Frazer and Co who lived in a house named “Shiel.” Proceeding further at No 36 was the home of Physician Dr VEP Seneviratne. Around here were the homes named Chetwynd and Donnington belonging to DF Peiris, built around the turn of the Twentieth century.
DF Peiris’s daughter, Maud, married Thomas Lambert Fernando, the grandfather of Dr Srilal Fernando, a joint author of this memoir. Donnington was later occupied by ARM Ameen, Consul for Egypt. Chetwynd was later owned by DF Peiris’ younger brother, the father of orthopaedic surgeon Dr Rienzie Peiris. Adjoining Donnington and located northwards was “Greylands” the home of Mudaliyar JCS Fonseka a stalwart of the Orchid Circle of Ceylon. At No 48 was the home of former Minister Montague Jayawicjkreme on whose large property many houses have since been constructed.
A few properties away is Sukasthan Gardens, a cluster of homes built on the grounds of the former stately home of Sir Ponnambalam Ramanathan named “Sukasthan”. It was inherited by Ramanathan’s daughter, Sundari, who eventually sold it. Gynaecologist Dr PR Thiagarajah lived in one of the houses that were built there. Another well known resident of Sukasthan Gardens was LS Boys, a Director of Gordon Frazer and Co who lived in a house named “Shiel.” Proceeding further at No 36 was the home of Physician Dr VEP Seneviratne. Around here were the homes named Chetwynd and Donnington belonging to DF Peiris, built around the turn of the Twentieth century.
DF Peiris’s daughter, Maud, married Thomas Lambert Fernando, the grandfather of Dr Srilal Fernando, a joint author of this memoir. Donnington was later occupied by ARM Ameen, Consul for Egypt. Chetwynd was later owned by DF Peiris’ younger brother, the father of orthopaedic surgeon Dr Rienzie Peiris. Adjoining Donnington and located northwards was “Greylands” the home of Mudaliyar JCS Fonseka a stalwart of the Orchid Circle of Ceylon. At No 48 was the home of former Minister Montague Jayawicjkreme on whose large property many houses have since been constructed.
Proceeding towards Borella on the left side of Ward Place are the two major government health care institutions the Victoria Memorial Eye Hospital and the Dental Institute. The Victoria Memorial Eye Hospital was built in honour of the Jubilee of Queen Victoria in 1897 and constructed in 1906. Designed by architect Edward Skinner in traditional Indo Sarasenic lines, it is characterised by its red brick façade and the many turrets of Sarasenic design. Further down the road is the Government run Dental Institute. The Dental Institute was set up in the 1930s with Dr W Balendra as its first Director. Dr Balendra himself was a resident of Ward place. Alongside was Volkaart gardens where homes of the Directors of Volkaart Brothers were located . Further on was the home “St Brycedale” of Dr Richie Caldera, Obstetrician in Charge of the De Soysa Maternity Home located on Regent Street running parallel to Ward Place. At No 53 were four homes built around the 1960s one of which was the home of Dr Chris Raffel.
A home in Ward Place and two eminent doctors, father, and son, also from Ward Place featured in a much publicised murder trial called the “Duff House Case” in the 1930s. White House in Ward Place was a large elegant home belonging to Solomon Seneviratne who was married to the sister of Sir Solomon Dia

s Bandaranaike. Solomon Seneviratne himself owned broad acres and his country home was situated on his coconut estate in Kotikawatte, Angoda. Solomon’s son Stephen was like the father educated at Royal College, and later at Cambridge University, where he qualified as a Barrister. He did not practice at the bar and spent his time managing the cattle farm which he inherited. He soon became a keen and enthusiastic cattle breeder with an expert knowledge of animal husbandry.
He married Lilian de Alwis, sister of Leo de Alwis, who was married to a daughter of Sir Solomon Dias Bandaranaike. Leo’s wife was a sister of the late Prime Minister SWRD Bandaranaike. The life of Stephen and Lilian was tumultuous. They had many quarrels regarding Stephen’s intention to sell his home, White House. The couple lived in Duff House at No 4. Bagatelle Road rented out at Rs 100 a month, a considerable sum as rent in the 1930s.
Lilian had a troubled pregnancy which ended with the birth of their only child Terrence. She did not have a warm relationship with the son as she blamed him for her difficult pregnancy. Lilian was found one day dead in the living room of the house having inhaled chloroform. The case tested the strength of the family relationships within the Bandaranaike extended family. Here was Sir Solomon’s brother-in-law’s son accused of the murder of Sir Solomon’s son-in-law’s sister. The police were notified and Lilian’s family, particularly her brother Leo de Alwis, was convinced that Stephen had forced his wife to inhale a lethal dose of chloroform.
Dr S C Paul who was a close friend of Sir Solomon gave expert medical evidence to support that contention, which was rejected by Stephen who said that his wife was depressed and could have inhaled chloroform which Stephen kept for his animal husbandry.
Stephen was however charged with the murder of his wife before Justice MT Akbar. Stephen’s defence was supported by the expert medical evidence of Dr SC Paul’s son Dr Milroy Paul. In his direction to the jury, Justice Akbar ignored aspects of evidence that would benefit the accused, and consequently, the accused was found guilty of murdering his wife and sentenced to death. This was in 1936 when there was no Court of Criminal Appeal, so the accused appealed to the Privy Council which overturned the judgment of Akbar and acquitted Stephen. The Privy Council also made some scathing observations on the findings of the trial judge which led to Akbar suffering depression and submitting his resignation from the bench. Finally, it seemed that the murder trial ended in the trial of the presiding judge!
There were two other older well known homes on Ward Place.. One was Chateau Jubillee occupied by Adrian St V Jayewardene, Supreme Court Judge, and brother of JR Jayewardene’s father EW Jayewardene. The other was Fairy Hall built in 1880 the original home of Dr Simon de Melho Aserappah and his wife Emily Wake. It was part of the large homestead on which 20 years later Rao Mahal and other homes were constructed by the family of Dr SC Paul who married Dr Aserappah’s daughter Dora.
Interior of the Dr PH Amerasinghe home designed by Architect Minnettte de Silva
The house 53/3 Ward Place designed by Geoffrey Bawa for Dr Chris Raffel was sold by Dr Chris and Carmel Raffel to Ajit Saravanamuttu who resided there until his death in 2006. Next door at No 55 was “Villa Mirelle” the home of Dr Percy Kulasinghe also situated on a large block which has since been subdivided with a new road named Kulasinghe Gardens hosting several houses. In the adjoining block at No 57 stands today the hotel Jetwing Colombo. Dr Kulasinghe was for many years a Director of the Ceylon Insurance Co founded and managed by fellow Ward Place resident Justin Kotelawela.
At No 61 was the home of lawyer FR de Saram and wife Miriam (nee Pieris) acclaimed aesthete and oriental dancer in an era when women were rarely seen on stage. Her elder son Rohan de Saram is the internationally famous cellist. The De Sarams engaged renowned architect Geoffrey Bawa to design a new additional home on the grounds now bearing No:61/6. Another Rohan, Rohan Perera at 57/2 and his brother Dr Hari Perera, Psychiatrist, the sons of the eminent lawyer HV Perera had their homes also in Ward Place.
At No:65 a house named “Taprobane “was the home of proprietary planter SR Muttiahpillai owner of the 1,250 acre Naluwella Group in Balangoda. His son M Rajendran managed the family properties in Balangoda until the initiation of Land Reform, and was awarded an MBE in recognition of his services to agriculture. The Muttiahpillai Caddillac in metallic blue colour was an ubiquitous feature of life in Ward Place in the 1950s. The passing of time and the demand for quality blocks of land has led to the breaking up of their large tract of land. A new road goes through the property now with the name Muththiahpillai Gardens, serving many new homes.
Dr W Balendra the dental surgeon’s home stood next door at No 67 next door to whom lived Dr May Ratnayake at No 69. Somewhere here stands the home of gynaecologist Dr PH (Chandra) Amerasinghe designed by renowned woman architect Minnette de Silva. She also designed the home of Chandra’s brother, Dr Asoka Amerasinghe in 5th Lane. Chandra was snatched away in his prime, from injuries resulting from an accident arising from a fun filled motor cycle ride.
The architect VS Thurairajah built a block of Flats at No 75 which was almost entirely leased out by the Marga Institute on its establishment in 1972. By 1975 Marga was in its own home at 61 Greenlands Avenue now known as Issipatana Mawata. Dr AC Arulpragasam ENT Surgeon and Dr Rajah Cooke both from the extended Paul family lived at No 77 as part of the large landholding adjacent to the Paul home “Rao Mahal “. Rao Mahal was built by Dr Simon De Melho Aserappah one of the first overseas qualified doctors who returned from England in the 19th Century. His daughter Dora married Dr SC Paul whose descendants still live in the original homestead in Ward Place where the
Paul family still retain a large extent of land on the site.
Dr Gunaratnam Cooke lived at 77 Ward Place, and Egerton Paul, another son of of Dr SC Paul, lived at No 85. Dr S.C Paul’s son, Dr Milroy Paul was the acclaimed surgeon who obtained his Master of Surgery qualification in the UK and was given the signal honour of delivering the “Hunterian Lecture” to the Royal College of Surgeons in England. He inherited Rao Mahal. Prof Milroy Paul’s son, Avinder, has collaborated in this present enterprise on homes in Ward Place and his knowledge and memory has helped us immensely in putting together this piece for the readers of The Ceylankan
Ward Place was closely associated with the development of the medical profession in Sri Lanka, and its early residential character was dominated by the medical profession. From the beginning therefore it was a highly gentrified area within the metropolis. Many successful doctors lived there, but they certainly would have had some unsuccessful medical adventures too, in addition to others whose lives were decreed not to go any further. They did not have to go far thereafter, the General Cemetery Kanatte also part of the former Borella estate, was nearby to provide them everlasting peace!
A cursory study of the residential features of this precinct would reveal that today it has lost that once dominant association with the medical profession. The street is located in one of the most sought after areas for dwellings today, and where large homes and gardens once stood, are large blocks of luxury apartments. Opulence still reigns however, and there is little doubt that Ward Place will continue to play host to a privileged few.
(This originally appeared in the Ceylankan)
Concluded
Features
Retirement age for judges: Innovation and policy
I. The Constitutional Context
Independence of the judiciary is, without question, an essential element of a functioning democracy. In recognition of this, ample provision is made in the highest law of our country, the Constitution, to engender an environment in which the courts are able to fulfil their public responsibility with total acceptance.
As part of this protective apparatus, judges of the Supreme Court and the Court of Appeal are assured of security of tenure by the provision that “they shall not be removed except by an order of the President made after an address of Parliament supported by a majority of the total number of members of Parliament, (including those not present), has been presented to the President for such removal on the ground of proved misbehaviour or incapacity”[Article 107(2)]. Since this assurance holds good for the entirety of tenure, it follows that the age of retirement should be defined with certainty. This is done by the Constitution itself by the provision that “the age of retirement of judges of the Supreme Court shall be 65 years and of judges of the Court of Appeal shall be 63 years”[Article 107(5)].
II. A Proposal for Reform
This provision has been in force ever since the commencement of the Constitution. Significant public interest, therefore, has been aroused by the lead story in a newspaper, Anidda of 13 March, that the government is proposing to extend the term of office of judges of the Supreme Court and the Court of Appeal by a period of two years.
This proposal, if indeed it reflects the thinking of the government, is deeply disturbing from the standpoint of policy, and gives rise to grave consequences. The courts operating at the apex of the judicial structure are called upon to do justice between citizens and also between the state and members of the public. It is an indispensable principle governing the administration of justice that not the slightest shadow of doubt should arise in the public mind regarding the absolute objectivity and impartiality with which the courts approach this task.
What is proposed, if the newspaper report is authentic, is to confer on judges of two particular courts, the Supreme Court and the Court of Appeal, a substantial benefit or advantage in the form of extension of their years of service. The question is whether the implications of this initiative are healthy for the administration of justice.
III. Governing Considerations of Policy
What is at stake is a principle intuitively identified as a pillar of justice.
Reflecting firm convictions, the legal antecedents reiterate the established position with remarkable emphasis. The classical exposition of the seminal standard is, of course, the pronouncement by Lord Hewart: “It is not merely of some importance, but is of fundamental importance that justice should not only be done, but should manifestly and undoubtedly be seen to be done”. (Rex v. Sussex Justices, ex parte McCarthy). The underlying principle is that perception is no less important than reality. The mere appearance of partiality has been held to vitiate proceedings: Dissanayake v. Kaleel. In particular, reasonableness of apprehension in the mind of the parties to litigation is critical: Ranjit Thakur v. Union of India, a reasonable likelihood of bias being necessarily fatal (Manak Lal v. Prem Chaud Singhvi).
The overriding factor is unshaken public confidence in the judiciary: State of West Bengal v. Shivananda Pathak. The decision must be “demonstrably” (Saleem Marsoof J.) fair. The Bar Association of Sri Lanka has rightly declared: “The authority of the judiciary ultimately depends on the trust reposed in it by the people, which is sustained only when justice is administered in a visibly fair manner”.
Credibility is paramount in this regard. “Justice has to be seen to be believed” (J.B. Morton). Legality of the outcome is not decisive; process is of equal consequence. Judicial decisions, then, must withstand public scrutiny, not merely legal technicality: Mark Fernando J. in the Jana Ghosha case. Conceived as continuing vitality of natural justice principles, these are integral to justice itself: Samarawickrema J. in Fernando v. Attorney General. Institutional integrity depends on eliminating even the appearance of partiality (Mandal Vikas Nigam Ltd. v. Girja Shankar Pant), and “open justice is the cornerstone of our judicial system”: (Sahara India Real Estate Corporation Ltd. v. SEBI).
IV. Practical Constraints
Apart from these compelling considerations of policy, there are practical aspects which call for serious consideration. The effect of the proposal is that, among all judges operating at different levels in the judicature of Sri Lanka, judges of the Supreme Court and the Court of Appeal only, to the exclusion of all other judges, are singled out as the beneficiaries of the proposal. An inevitable result is that High Court and District Judges and Magistrates will find their avenues of promotion seriously impeded by the unexpected lengthening of the periods of service of currently serving judges in the two apex courts. Consequently, they will be required to retire at a point of time appreciably earlier than they had anticipated to relinquish judicial office because the prospect of promotion to higher courts, entailing higher age limits for retirement, is precipitately withdrawn. Some degree of demotivation, arising from denial of legitimate expectation, is therefore to be expected.
A possible response to this obvious problem is a decision to make the two-year extension applicable to all judicial officers, rather than confining it to judges of the two highest courts. This would solve the problem of disillusionment at lower levels of the judiciary, but other issues, clearly serious in their impact, will naturally arise.
Public service structures, to be equitable and effective, must be founded on principles of non-discrimination in respect of service conditions and related matters. Arbitrary or invidious treatment is destructive of this purpose. In determining the age of retirement of judges of the Supreme Court and the Court of Appeal, some attention has been properly paid to balance and consistency. The age of retirement of a Supreme Court judge is on par with that applicable to university professors and academic staff in the higher education system. They all retire at 65 years. Members of the public service, generally, retire at 60. Medical specialists retire at 63, with the possibility of extension in special circumstances to 65. The age of retirement for High Court Judges is 61, and for Magistrates and District Judges 60. It may be noted that the policy change in 2022 aimed at specifically addressing the issue of uniformity and compatibility.
If, then, an attempt is made to carve out an ad hoc principle strictly limited to judicial officers, not admitting of a self-evident rationale, the question would inevitably arise whether this is fair by other categories of the public service and whether the latter would not entertain a justifiable sense of grievance.
This is not merely a moral or ethical issue relating to motivation and fulfillment within the public service, but it could potentially give rise to critical legal issues. It is certainly arguable that the proposed course of action represents an infringement of the postulate of equality of treatment, and non-discrimination, enshrined in Article 12(1) of the Constitution.
There would, as well, be the awkward situation that this issue, almost certain to be raised, would then have to be adjudicated upon by the Supreme Court, itself the direct and exclusive beneficiary of the impugned measure.
V. Piecemeal Amendment or an Overall Approach?
If innovation on these lines is contemplated, would it not be desirable to take up the issue as part of the new Constitution, which the government has pledged to formulate and enact, rather than as a piecemeal amendment at this moment to the existing Constitution? After all, Chapter XV, dealing with the Judiciary, contains provisions interlinked with other salient features of the Constitution, and an integrated approach would seem preferable.
VI. Conclusion
In sum, then, it is submitted that the proposed change is injurious to the institutional integrity of the judiciary and to the prestige and stature of judges, and that it should not be implemented without full consideration of all the issues involved.
By Professor G. L. Peiris
D. Phil. (Oxford), Ph. D. (Sri Lanka);
Former Minister of Justice, Constitutional Affairs and National Integration;
Quondam Visiting Fellow of the Universities of Oxford, Cambridge and London;
Former Vice-Chancellor and Emeritus Professor of Law of the University of Colombo.
Features
Ranked 134th in Happiness: Rethinking Sri Lanka’s development through happiness, youth wellbeing and resilience
In recent years, Sri Lanka has experienced a succession of overlapping challenges that have tested its resilience. Cyclone Ditwah struck Sri Lanka in November last year, significantly disrupting the normal lives of its citizens. The infrastructure damage is much more serious than the tsunami. According to World Bank reports and preliminary estimates, the losses amounted to approximately US$ 4.1 billion, nearly 4 per cent of the country’s Gross Domestic Product. Before taking a break from that, the emerging crisis in the Middle East has once again raised concerns about potential economic repercussions. In particular, those already affected by disasters such as Cyclone Ditwah risk falling “from the frying pan into the fire,” facing multiple hardships simultaneously. Currently, we see fuel prices rising, four-day workweeks, a higher cost of living, increased pressure on household incomes, and a reduction in the overall standard of living for ordinary citizens. It would certainly affect people’s happiness. As human beings, we naturally aspire to live happy and fulfilling lives. At a time when the world is increasingly talking about happiness and wellbeing, the World Happiness Report provides a useful way of looking at how countries are doing. The World Happiness Report discusses global well-being and offers strategies to improve it. The report is produced annually with contributions from the University of Oxford’s Wellbeing Research Centre, Gallup, the UN Sustainable Development Solutions Network, and other stakeholders. There are many variables taken into consideration for the index, including the core measure (Cantril Ladder) and six explanatory variables (GDP per Capita ,Social Support,Healthy Life Expectancy,Freedom to Make Life Choices,Generosity,Perceptions of Corruption), with a final comparison.
According to the recently published World Happiness Report 2026, Sri Lanka ranks 134th out of 147 nations. As per the report, this is the first time that Sri Lanka has suffered such a decline. Sri Lanka currently trails behind most of its South Asian neighbours in the happiness index. The World Happiness Report 2026 attributes Sri Lanka’s low ranking (134th) to a combination of persistent economic struggles, social challenges, and modern pressures on younger generations. The 2026 report specifically noted that excessive social media use is a growing factor contributing to declining life satisfaction among young people globally, including in Sri Lanka. This calls for greater vigilance and careful reflection. These concerns should be examined alongside key observations, particularly in the context of education reforms in Sri Lanka, which must look beyond their immediate scope and engage more meaningfully with the country’s future.
In recent years, a series of events has triggered political upheaval in countries such as Nepal, characterised by widespread protests, government collapse, and the emergence of interim administration. Most reports and news outlets described this as “Gen Z protests.” First, we need to understand what Generation Z is and its key attributes. Born between 1997 and 2012, Generation Z represents the first truly “digital native” generation—raised not just with the internet, but immersed in it. Their lives revolve around digital ecosystems: TikTok sets cultural trends, Instagram fuels discovery, YouTube delivers learning, and WhatsApp sustains peer communities. This constant, feed-driven engagement shapes not only how they consume content but how they think, act, and spend. Tech-savvy and socially aware, Gen Z holds brands to a higher standard. For them, authenticity, transparency, and accountability—especially on environmental and ethical issues—aren’t marketing tools; they’re baseline expectations. We can also observe instances of them becoming unnecessarily arrogant in making quick decisions and becoming tools of some harmful anti-social ideological groups. However, we must understand that any generation should have proper education about certain aspects of the normal world, such as respecting others, listening to others, and living well. More interestingly, a global survey by the McKinsey Health Institute, covering 42,083 people across 26 countries, finds that Gen Z reports poorer mental health than older cohorts and is more likely to perceive social media as harmful.
Youth health behaviour in Sri Lanka reveals growing concerns in mental health and wellbeing. Around 18% of youth (here, school-going adolescents aged 13-17) experience depression, 22.4% feel lonely, and 11.9% struggle with sleep due to worry, with issues rising alongside digital exposure. Suicide-related risks are significant, with notable proportions reporting thoughts, plans, and attempts, particularly among females. Bullying remains a significant concern, particularly among males, with cyberbullying emerging as a notable issue. At the same time, substance use is increasing, including tobacco, smokeless tobacco, and e-cigarettes. These trends highlight the urgent need for targeted interventions to support youth mental health, resilience, and healthier behavioural outcomes in Sri Lanka. We need to create a forum in Sri Lanka to keep young people informed about this. Sri Lanka can designate a date (like April 25th) as a National Youth Empowerment Day to strengthen youth mental health and suicide prevention efforts. This should be supported by a comprehensive, multi-sectoral strategy aligned with basic global guidelines. Key priorities include school-based emotional learning, counselling services, and mental health training for teachers and parents. Strengthening data systems, reducing access to harmful means, and promoting responsible media reporting are essential. Empowering families and communities through awareness and digital tools will ensure this day becomes a meaningful national call to action.
As discussed earlier, Sri Lanka must carefully understand and respond to the challenges arising from its ongoing changes. Sri Lanka should establish an immediate task force comprising responsible stakeholders to engage in discussions on ongoing concerns. Recognising that it is not a comprehensive solution, the World Happiness Index can nevertheless act as an important indicator in guiding a paradigm shift in how we approach education and economic development. For a country seeking to reposition itself globally, Sri Lanka must adopt stronger, more effective strategies across multiple sectors. Building a resilient and prosperous future requires sound policymaking and clear strategic direction.
(The writer is a Professor in Management Studies at the Open University of Sri Lanka. You can reach Professor Abeysekera via nabey@ou.ac.lk)
by Prof. Nalin Abeysekera
Features
Hidden diversity in Sri Lanka’s killifish revealed: New study reshapes understanding of island’s freshwater biodiversity
A groundbreaking new study led by an international team of scientists, including Sri Lankan researcher Tharindu Ranasinghe, has uncovered striking genetic distinctions in two closely related killifish species—reshaping long-standing assumptions about freshwater biodiversity shared between Sri Lanka and India.
Published recently in Zootaxa, the research brings together leading ichthyologists such as Hiranya Sudasinghe, Madhava Meegaskumbura, Neelesh Dahanukar and Rajeev Raghavan, alongside other regional experts, highlighting a growing South Asian collaboration in biodiversity science.
For decades, scientists debated whether Aplocheilus blockii and Aplocheilus parvus were in fact the same species. But the new genetic analysis confirms they are “distinct, reciprocally monophyletic sister species,” providing long-awaited clarity to their taxonomic identity.
Speaking to The Island, Ranasinghe said the findings underscore the hidden complexity of Sri Lanka’s freshwater ecosystems.
“What appears superficially similar can be genetically very different,” he noted. “Our study shows that even widespread, common-looking species can hold deep evolutionary histories that we are only now beginning to understand.”
A tale of two fishes
The study reveals that Aplocheilus blockii is restricted to peninsular India, while Aplocheilus parvus occurs both in southern India and across Sri Lanka’s lowland wetlands.
Despite their close relationship, the two species show clear genetic separation, with a measurable “genetic gap” distinguishing them. Subtle physical differences—such as the pattern of iridescent scales—also help scientists tell them apart.
Co-author Sudasinghe, who has led several landmark studies on Sri Lankan freshwater fishes, noted that such integrative approaches combining genetics and morphology are redefining taxonomy in the region.
Echoes of ancient land bridges
The findings also shed light on the ancient biogeographic links between Sri Lanka and India.
Scientists believe that during periods of low sea levels in the past, the two landmasses were connected by the now-submerged Palk Isthmus, allowing freshwater species to move between them.
Later, rising seas severed this connection, isolating populations and driving genetic divergence.
“These fishes likely dispersed between India and Sri Lanka when the land bridge existed,” Ranasinghe said. “Subsequent isolation has resulted in the patterns of genetic structure we see today.”
Meegaskumbura emphasised that such patterns are increasingly being observed across multiple freshwater fish groups in Sri Lanka, pointing to a shared evolutionary history shaped by geography and climate.
A deeper genetic divide
One of the study’s most striking findings is that Sri Lankan populations of A. parvus are genetically distinct from those in India, with no shared haplotypes between the two regions.
Dahanukar explained that this level of differentiation, despite relatively recent geological separation, highlights how quickly freshwater species can diverge when isolated.
Meanwhile, Raghavan pointed out that these findings reinforce the importance of conserving habitats across both countries, as each region harbours unique genetic diversity.
Implications for conservation
The study carries important implications for conservation, particularly in a country like Sri Lanka where freshwater ecosystems are under increasing pressure from development, pollution, and climate change.
Ranasinghe stressed that understanding genetic diversity is key to protecting species effectively.
“If we treat all populations as identical, we risk losing unique genetic lineages,” he warned. “Conservation planning must recognise these hidden differences.”
Sri Lanka is already recognised as a global biodiversity hotspot, but studies like this suggest that its biological richness may be even greater than previously thought.
A broader scientific shift
The research also contributes to a growing body of work by scientists such as Sudasinghe and Meegaskumbura, challenging traditional assumptions about species distributions in the region.
Earlier studies often assumed that many freshwater fish species were shared uniformly between India and Sri Lanka. However, modern genetic tools are revealing a far more complex picture—one shaped by ancient geography, climatic shifts, and evolutionary processes.
“We are moving from a simplistic view of biodiversity to a much more nuanced understanding,” Ranasinghe said. “And Sri Lanka is proving to be a fascinating natural laboratory for this kind of research.”
Looking ahead
The researchers emphasise that much remains to be explored, with several freshwater fish groups in Sri Lanka still poorly understood at the genetic level.
For Sri Lanka, the message is clear: beneath its rivers, tanks, and wetlands lies a largely untapped reservoir of evolutionary history.
As Ranasinghe puts it:
“Every stream could hold a story of millions of years in the making. We are only just beginning to read them.”
By Ifham Nizam
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